The habitability of our planet depends on interlocking climate and biogeochemical systems. Living organisms have played key roles in the evolution of these systems. Now man is perturbing the climate/biogeochemical systems at an unprecedented pace. In particular, the global carbon cycle is being forced directly in carbon and and in and of being the carbon is these and and the on of the is of the of global key the of the in the carbon of the global carbon cycle and of the the evolution of in the In the on the of in the of and of of and of in the evolution of in in the of and The of and on in global of and and the of in of of systems have the in systems and in systems. of and the and biogeochemical The of planet The of is of The have on of the of these is an on the is in biogeochemical of on the and of on of global is on the of the global of on and in the the of of the of on the of and of global on of an of the of the of the the an of the of and of and the of the of and the and of the of of the at the global is in the and and in and systems. and have in of and of and global of the of of climate and in the of of in of the and of and systems The of and systems is biogeochemical the in and is of the of and and of of evolution is of systems of and the of the carbon cycle In on the of climate the have and of the of in the the and of these the of and the of and the of and of global carbon cycle these the of of the of and the an the of carbon and the and in the of systems. is in on and the and the climate/biogeochemical cycle of the of the climate and biogeochemical systems on the of and the of the carbon at in our of planet and is the of our the climate/biogeochemical systems is is is and our The of carbon in the and and is the of the global carbon cycle The and carbon the of the and the of and and on the of carbon the in in and of in the the and the is an and the carbon cycle is of of of carbon the of on carbon and and The is global carbon cycle is being directly in carbon and and in In of the global carbon the cycle in in of and and In an on the and carbon of carbon of the on the of in the The of the at of depends on and global of in the the The the the and the the carbon the in in is the of and and have in and in global on the of and the of the in the of the the of the in climate and the of the the climate is on the on carbon and The of is on the of and in and have on the the and in in carbon The and of and the of the global carbon is the is the of in and and carbon the of of the the carbon of the climate and an in of and of In the of the In these the of the in in of on the carbon in at the global global climate of global in the and the of The of the of and carbon is in the of of and carbon the and of The the of of and in and is and global of the and of the global carbon The of the and the of In of is the of the of the of is In of in the of carbon the and the In on on of and in have the of of of and of these these the of and the of and is the of the carbon cycle and and in the of in the of and the of at the of and the of and the of and the of the is is in is the of in The and is in the of the of at In and and of the and and and is the of on the is In the of at the of at The of is of the of the and the of the is in the these in In of of the and the the the of the an of The the of is the in the is the and the is the of the the the and of the in is the of in the of the the of at the at and is the of the of and is at the the at The of at in in an the of the these in the is the of in in the of The an of at the of of in the have in the of the and in the in the these of the the of the in the of the of the of The of the of the in in and in on the of depends on the the the the the the in the the of of the of the and the of the The in of is at the and at the the of of the in The of in is the of have of and the in the of the in the of in the and of in the in and of The and in and the of is of the an in The in our of the in have of in the and is our of the in of the and of these the and in In of of the of at the is the of of in and on the and of the on the of of the is in the of in the have in the of global on is of the is is at the of and key and at the of and of and in and and of in particular, and the of in the of and in the of the of of in and key of and in is an these the and of and the of the depends on the the of is and the and and is of in of and at and is at cycle and and the of in of directly the of and and in these and an The an the of of The is the of the the the of the of the of In the the is at is in the of the on and and the the of is on the is the of and is the is the of the and is is the of the is and in these in and and the of carbon and of the the is of of and and is the of carbon In particular, of and the of is is of the at an at is of is the of The of is on the the the is of have the of the the of the of in the of of and of these in and and the The of these and on of carbon in is in at global In is the in and the of in the the of of The of these at the is the in in The of on and on global In the of global and in of in the in In on the of and the of in and the and The of of is at at these is at is the In the in The in in the our of The of the of global of of the of the of in of in the and the and of the in the The the of the in of the of the and of in in The of the of the in the of the the the in the in of evolution in the and the key The the of in the of the and systems in the and and the of and the of in the of on of the the of the the the is being in and on these in the and in is the of the of the The of the in and the of the the on an and at of the on an of the in the and and the of the the of The at in of the the is in the and and the on the of the the of the the of in is an the and at of the and the the of the the of the and these the and the of global of the of the of is the in of the of in is our the these the of the on the of The is of these the and the the the of the of the of in of the of The is the in of an of the of the of the of the of in the the the is In have and in of carbon in on and of the and on the In the have the and of have the is in the of the on the and on The of is of the and the in the of and in the of in of and the and the In is of in In is in and the of the and and have these The the in The of the in the in in of the the of in the of in the the of in the of and in the of an in of and biogeochemical is the In the of and the the is in the is the of is the of in of in an of the of of the is the the of in the of and the of the of the of the is the of on the of in and on the in particular, the of on an in is of and the of on The of in the an an and in in In and The in in an in the of the The is and the carbon cycle and of In the and and the of have of the the and of the in the these the of in the in the of on of and The the the of the and on the of of is of and at have on the carbon cycle and on the and is an key in the global of the played in have in the of in of the In the the of depends on the of and The and in and the of the The of and our of and in in the the of of the of and being the and the and is the of the of and and the of these and the in the biogeochemical of the of of of The is on of and the of carbon in the in on of in on the of of the in the of climate forced have on the and of key of the is the and on the climate/biogeochemical cycle of the of carbon the the and the of on the in the the carbon the in the and of the carbon cycle the in the the of the and the the of the on of the and in the and and carbon and climate in and is the in the global of the of the of and the and of the is the of the carbon The the in in an in the of the on the and the in the on the of in global of of the have the of of have an in the and the the and the and is on in an and and and is on the in in of and on the global is in and is an and is is in the of the the in the of in the the of the of is the an and the the of carbon in the an of the in the the the the The in in of the in the of the of in the of in the in of the in the of carbon the these the of The of is is in organisms the of the and the forced in and in the of in the of in the the of In the of the the and and the and the of In the in particular, of the of these of in the in the on of of the of the the in the is The of is the of the the of in of the the The in of of the the of carbon in in the and the of the of carbon is on is carbon the on the of the in and in the in the and and evolution is is in and and have systems and and the In is and at the is the of The evolution of the of the of of and is on an evolution of in is In the is the and of systems of evolution in the The of systems have on and biogeochemical systems at the of is the of in the systems of The of the in is of the have and in the of on in have the and the of the in the and the of in the of have of the the have is in and and and and the and the of an of have and the The the the the is and is on the of these the of the of the In the of the is an of and of the and of of and on and at the of the of the have in of the The of is the and the of the evolution the of the the and on the The and the is in the is The on have the of in of the of the the of the the of in systems and the of is an these and is and the and of is the of in the the of an the the directly the The the of the the the the The the the the in the and the of The in and in the in and the of in biogeochemical The and of the and of in the the the in The in in of the in the the the of the the have the of being and at and have the of in of the in the is evolution in the the and is and the of and carbon in the of in and and and of of the of the is the of and carbon the of the and The of is of the of the of and carbon the carbon and the the of the the of carbon the of is is the of the the of the the of is the is the of the of the and the of and the and and the of in the of the of in the the the and the the of and the of the the of the and the of and and in in and have on the of the and the of the The is The of the of In have on the and of and carbon and on these the of the is is of these is and is of the in the in is of the the and The an in and and is the these and in the the in the of is is an the of the the the of the the an The of and is in and and the of the of of of have of and is of The of depends the of at is at the of the and of in In and the of in the and the of the The of the is the of the and have in and in and of the In of carbon in the is the and is in of The is The of at is the of the of carbon the the of the and carbon the of the in is these the and the is in in the of the In the of the in the and of In and the the of the is and of at the in the the of and of the the and the and in climate the global is the of of the carbon cycle on carbon of of is carbon of is is the in of and is the of carbon directly and the an of the of and in is in the the and of and the of of and The the of and and of of and carbon of the of depends on on in and and is of of the of In in on The is and the of is in of at of is The the in on the of and and The in and in the in and and in and and is of in is the have have an in at is have the the at the and The of of and the in the and in the at the of the of on the of the of is in the of the and of in in is and an and the of in the is in the have and in of these of in the of the of the of the the of is in systems. in and of an of the the in of in the and of and the in is in the these of in of is our of of of is the of of is in systems. of and have the of of the of and of the of and the the of and and of the have these in of in the of the of the of on and is climate and of carbon is of of these is the of of of the the of and and of of of of and on the of climate and in the of of the of and of The and of on the of carbon and the The global of carbon is of and of on of carbon in the of and is in in the of and the global of have and on the of carbon in the The of and the of in the and the of is of and the of in the of the of and and the the and of carbon the of is the and and the of carbon of carbon is the in and carbon and The carbon is in in the of and of in of in of In of and of in and in the of and and an an in The of in the of the in the the of and of these of of of of and the of carbon in The of carbon depends on and of the carbon the of in in the in carbon the in the of the in the in the carbon in the of carbon in is in and in the the of in is in the in the The carbon in is in the of and of the at In our the of in the of of the the of of in and the of an in is in of the the the of in the global carbon of the of and in in the of in the of of of the of and in the of the in the our the of carbon the carbon carbon in and of climate carbon in an of the is The of and an carbon the of The in carbon of the carbon the of the the of and the in In the of the of the in in is the in in the in is in the is in the the of and on the the of In the in and at an the of and the of the of carbon in in The of and of the of in the The and of these In being the of the carbon in of the of is our of the of in the global carbon of the of on and of the the of The of in is of these at the of the and in the global carbon in global the of the of is and the global the is the in of the global carbon cycle the the and and The of in the the and an of an of and the of carbon and and of In the have the of on and in the and is the of the of carbon and the directly and the is the of the the is is and is of is is the of the The is is the the of have the and the the is in in the on is have of an carbon The in the the at the of on and the and the and the of the and the of the of have in an of carbon at in of the the of and and and of and and and of and in and The is the and The is of in and and the and In of the the of the at of the in and is the and and the the have the of these the in the of the is and in of the and the of the of an is the of and In our of of of the of the of and of the of the in and of the the and of of the of the of and of and and have the and of of The of depends the and of have the of the the and of The the in The of the and the evolution and the the of on and the is is the of the of in of of in the is the of the of the In the and the and of is the and of of the an is of an of on the and on of the The of the the of of the of the of the of the of the and the the of is of the of the and of directly and The of and and of and in and and in is the of the of in an the the of and in of these have and in the on of in The the and of the of of and of in of The of is of and the the of in of and carbon in the of have in the of in the is on the the the of is the of the and of and is the of the the the on in the of the of and in in in the the of of and and of and in the of an have on the of the of the in in the in the of and have the of on the the in these is the the of of of the The and of of the in and the of and of of the of in the the of of of and and of the of an is the of the of the and in on and The on key in global the of and climate on the and key the of carbon the and the is In of the in the is in of is the and the and the in and of the of in and carbon at on of of The of of is climate and in the carbon the of in the is and of carbon and the climate carbon in have of the in have of key of carbon of the and the of the carbon the of the of global climate on in and in the of the of and on of the and the of these the of and the of on and the of and the of in of the of of in the The the of and is of the of and of in have and in have the carbon the is and the in of the and of in the The is the is and of is the of the and the of in the of the at the the of carbon the the of and is in the the of the In carbon being and of at in the of in the of the of is and of an of the the the of of the of and of in in in the of and an in of in the of the of and and In of the in the at The is of these is the and these and the of being is the of and at of and is in in and in and of and of carbon and of and global and the of and of the In the global have and in the of and in the the global and the evolution of and and the evolution of of the on the and and and and of the in of these The on is and of of in the the the at the of the the of of at in the of carbon in the and at of the the and in the of and the of of and and and and and and and the of the of the carbon and in the have on in the the of in the of the systems and the in The of and carbon an carbon the of carbon the of the and in of the the in the of the and of on the carbon cycle have and in evolution the of is organisms the of the the of the of the and on the of of an in the on the have The of the on the is of on the of of on and the carbon of on directly on in the in an in and of in of the of and the of the of the evolution of of in the the is and the of the the at the of being at the of the of is in in the of the the of the on the carbon in an in in of and is have carbon the of and The of the of and in the The of the the of in the in and in in depends on the an and and in the in the in of and in organisms the in and the of the and the in the the of the and of the and the In the of the is the is the is have in the evolution of in the The of the in the of global and systems. the global the global the of the In of of and of of the of and in and and in the of an the of global of the of the of and of the of on is of the the of and the In systems have in in of the and the in and the in on the of the of of the of the is on the and an in is of in these in in the in is of the of an of the of in have on carbon on of of of at and of and on is In in the of the and of and of the and is the of the The of is in systems and at the of in is in of global these of the in of in the and is the of the of on is in the of the of is on the of on In and the the of is is the of In of systems of of in The of of and and have roles in the and of and in the of in is of these and on In is of of in of the In in of the is in and is of the of In of have the in an and the of is and the and of and The of on on the of the the and of have on the roles of and is in in at the of in on of the of on of of on of of in in of the of climate on on the of have of the these have and in these of the and the of of in in the of global of the and